the understorey compositional dissimilarity between patches within a stand increases with increasing number of tree species in the overstorey, while the compositional dissimilarity between different stands decreases with higher tree species richness

Comparatively homogeneous environmental conditions in monocultures contrast with the heterogeneous pattern of patches with distinct resource availability and soil conditions within mixed stands (Morin et al., 2011; Yankelevich et al., 2006). Each understorey species has its own optimal requirements concerning resources and soil conditions. The relatively higher environmental heterogeneity within a mixed stand might thus be reflected in an elevated compositional dissimilarity between patches (Golodets et al., 2011). The more homogeneous distribution of resources and soil conditions in monocultures are mirrored in a uniform understorey composition (Beatty, 2003). At the same time, resource availability and soil conditions in monocultures can reasonably be assumed to be quite divergent from monocultures or mixtures with other tree species or species that are not closely related. As mixed stands within a certain region often share tree species, or species with a similar impact on resources and soil conditions, with monocultures and other mixed stands, the dissimilarity in resources and soil conditions with other stands is expected to be lower compared to monocultures.

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